The Impacts of
Mountain Biking on Wildlife and People --
A Review of the
Literature
Michael J.
Vandeman, Ph.D.
July 3, 2004
"Every recreationist -- whether hiker, biker, horsepacker, or posey sniffer --
should not begin by asking, 'What's best for ME?' but rather 'What's best for
the bears?'" Tom Butler
"Will we keep some parts of the American landscape
natural and wild and free -- or must every acre be easily accessible to people
and their toys? … Mountain bikes' impacts on the land are large and getting
worse. … The aggressive push of mountain bike organizations to build
ever-growing webs of trails poses serious problems of habitat fragmentation,
increased erosion, and wildlife conflicts.
As interest in
extreme riding continues to grow, as trail networks burgeon, and as new
technology makes it possible for ever-more mountain bicyclists to participate,
even the most remote wild landscapes may become trammeled -- and trampled -- by
knobby tires. … The destruction of wilderness and the fragmentation of habitats
and ecosystems is death by a thousand cuts. Will introduction of mountain bikes
-- and their penetration farther into wilderness -- promote additional
fragmentation and human conflicts with the natural world? Yes." Brian
O'Donnell and Michael Carroll
"Some things are obvious: mountain bikes do more
damage to the land than hikers. To think otherwise ignores the story told by
the ground. Although I have never ridden a mountain bike, I am very familiar
with their impacts. For the last seven years I have regularly run three to six
miles several times a week on a network of trails in the
Because I'm
clumsy, I keep my eyes on the trail in front of me. I run or walk in all
seasons, in all kinds of weather. I have watched the growing erosion on these
trails from mountain bike use. The basic difference between feet and tires is
that tire tracks are continuous and foot tracks are discontinuous. Water finds
that narrow, continuous tire tracks are a rill in which to flow. Also, because
many mountain bikers are after thrills and speed, their tires cut into the
ground. Slamming on the brakes after zooming downhill,
sliding around sharp corners, and digging in to go uphill: I see the results of
this behavior weekly. …
I regularly see
mountain bikers cutting off cross-country, even on steep slopes, for more of a
challenge. They seem blind and deaf to the damage they cause. Admittedly,
backpackers and horsepackers can cause damage to
wilderness trails. But this is a poor argument to suggest that we add another
source of damage to those trails." Dave Foreman
"Studies show that bike impacts are similar to those of other
non-motorized trail users." Jim Hasenauer
(professor of rhetoric and member of the board of directors of the
International Mountain Bicyclists Association)
Introduction:
I
first became interested in the problem of mountain biking in 1994. I had been
studying the impacts of the presence of humans on wildlife, and had come to the
conclusion that there needs to be habitat that is entirely off-limits to
humans, in order that wildlife that is sensitive to the presence of humans can
survive (see Vandeman, 2000). But what is the best way to minimize the presence
of people? Restricting human access is repugnant, and difficult and expensive
to accomplish. It occurred to me that the best way to reduce the presence and
impacts of humans is to restrict the technologies that they are allowed
to utilize in nature: e.g. prohibit bicycles and other vehicles (and perhaps
even domesticated animals, when used as vehicles).
Having
been a transportation activist for eight years (working on stopping highway
construction), and having a favorable view of my fellow bicyclists as
environmentalists, I turned to them to help me campaign to keep bicycles out of
natural areas. Was I ever surprised! I discovered that many bicyclists (e.g.
many mountain bikers) aren't environmentalists at all, but are simply people
who like to bicycle -- in the case of mountain bikers,
many of them just use nature, as a kind of playground or outdoor
gymnasium! (Of course, there are also hikers, equestrians, and other
recreationists who fall into this category.) To my suggestion to keep bikes off
of trails in order to protect wildlife, they reacted with hostility! (There is
a degree of balkanization among activists, where some transportation activists
ignore the needs of wildlife, and some wildlife activists eschew bikes and
public transit.)
In
1994 I attended a public hearing held by the East Bay Municipal Utility (water)
District to decide whether to allow bikes on their watershed lands. Mountain
bikers were there asking for bike access, and the Sierra Club was there to
retain the right to hike, while keeping out the bicycles. I said that I had no
interest in using the watershed, but that I wanted to ensure that the wildlife
are protected -- hence, I asked that bikes not be allowed. Afterward, the EBMUD
Board of Directors took a field trip to
It
is obvious that mountain biking is harmful to some wildlife and people. No one,
even mountain bikers, tries to deny that. Bikes create V-shaped ruts in trails,
throw dirt to the outside on turns, crush small plants and animals on and under
the trail, facilitate increased levels of human access into wildlife habitat, and
drive other trail users (many of whom are seeking the tranquility and
primitiveness of natural surroundings) out of the parks. Because land managers
were starting to ban bikes from trails, the mountain bikers decided to try to
shift the battlefield to science, and try to convince people that mountain
biking is no more harmful than hiking. But there are two problems with this
approach: (1) it's not true, and (2) it's irrelevant.
I
will examine (1) in a moment. But first, let's look at relevance: whether or
not hiking (or All Terrain Vehicles or urban sprawl or anything else) is
harmful really has no bearing on whether mountain biking is harmful: they are
independent questions. Such a comparison would only be relevant if one were
committed to allowing only one activity or the other, and wanted to know which
is more harmful. In reality, hiking is always allowed, and the question is
whether to add mountain biking as a permitted activity. In that case,
the only relevant question is: Is mountain biking harmful? Of course, it is.
However, since many people seem interested in the outcome of the comparison, I
will examine the research and try to answer it.
The
mountain bikers' other line of research aims to prove that mountain bikers are
just like hikers, implying that they should have the same privileges as hikers.
(Of course, they already have the same privileges! The exact same rules
apply to both groups: both are allowed to hike everywhere, and neither is
allowed to bring a bike where they aren't allowed.) Using surveys, they have
tried to show that mountain bikers are really environmentalists, lovers of
nature, and deep ecologists. Of course, surveys are notoriously unreliable:
statements of belief don't easily translate into behavior. I'm going to ignore
this research, since I am (and the wildlife are) more interested in actual
impacts, not intentions.
The
International Mountain Biking Association (IMBA) has done me the favor of
collecting all the research they could find that seemed favorable to mountain
biking. Gary Sprung (2004) summarized it in his carefully worded essay,
"Natural Resource Impacts of Mountain Biking".
Impacts on Soil (Erosion):
This
study is frequently cited by mountain bikers as proof that mountain biking
doesn't cause more impact than hiking. But it has a number of defects that call
its conclusions into question. The authors used a "rainfall
simulator" to measure "sediment made available" by the various
treatments. They "[collected] surface runoff and sediment yield produced by
the simulated rainstorms at the downslope end of each
plot", which they claim "correlates with erosion" (they don't
say what the correlation coefficient is). This doesn't seem like a good measure
of erosion. For example, if a large rock were dislodged, the very weak
"simulated rainfall" wouldn't be capable of transporting it into the
collecting tray; only very fine particles would be collected. In fact, they
admit that the simulator's "small size … meant that the kinetic energy of
the simulated rainfall events was roughly one-third that of natural
rainstorms". Another reason to suspect that the measurements aren't valid
is that "none of the relationships between water runoff and soil texture,
slope, antecedent soil moisture, trail roughness, and soil resistance was statistically
significant".
Another
problem with the study is that the hikers and mountain bikers used trails that
were significantly different, prior to the experiment!: "The results from
Part A of Table 4 suggest that the trails used for the five treatment types
were not similar in terms of their sediment yield behavior prior to the
treatments. Trail plots used for hikers were statistically different from one
of the other groups (off-road bicycles) at the .05 level" (p.84). This
makes it even less likely that the hiker-mountain biker comparison is valid.
The
authors also ignored the relative distances that various trail users typically
travel (for example, bikers generally travel several times as far as hikers,
multiplying their impacts accordingly) and the additional impacts due to the
mountain bike bringing new people to the trails that otherwise would not have
been there (the same omission is true of all other studies, except Wisdom et al
(2004)). They do say "Trail use in the last ten years has seen a dramatic
increase in off-road bicycles" (p.86), but they don't incorporate this
fact into their comparison. In addition, there is no recognition of different
styles of riding and their effect on erosion. We don't know if the mountain
bikers rode in representative fashion, or, more likely, rode more gently, with
less skidding, acceleration, braking, and turning. There was also no
recognition that soil displaced sideways (rather than downhill) also
constitutes erosion damage. It seems likely that they underestimated the true
impacts of mountain biking. I don't think that these results are reliable.
(Note that the study was partially funded by IMBA.)
Besides
ignoring distance travelled, there were a number of other defects in the study.
The biking that was compared with hiking was apparently not typical mountain
biking. It was apparently slower than normal and included no skidding. Bikers
who skidded (a normal occurrence) were not compared with hikers. Their erosion
impacts were much greater than those of any hikers (judging from the study's
graph labelled "Figure 3"). Bikers' impacts
under wet conditions were also greater than those of the hikers, which probably
would have been statistically significant, if the numbers (of data points) had
been greater. One useful result was that the bikers tended to create a V-shaped
groove, whereas the hikers' impact was spread more evenly across the trail.
They admit that this "could act as a water
channel and increase erosion" (p.356). They also surveyed trail users:
"34% of riders listed excitement/risk as a main reason for visiting [the
park]. This, combined with the 57% of 'other users' who visit for relaxation,
sets up a potential for goal interference, in that a rider aiming for an
exciting/risky experience has the potential to interfere with a walker aiming
to have a relaxing experience." (p.357) This
would also tend to indicate that many bikers travel faster than those in this
study, since they are seeking "excitement" and "risk".
Impacts on Plants:
He
cites Thurston and Reader (2001), claiming that "hiking and bicycling
trample vegetation at equal rates … the impacts of biking and hiking measured
here were not significantly different". Actually, that is not true.
Although overall impacts weren't significantly different, "soil exposure
[was] greater on biking 500 pass lanes than hiking 500 pass lanes"
(p.404). In other words, after 500 passes, mountain biking began to show
significantly greater impacts. Thus their conclusion, "the impacts of
biking and hiking measured here were not significantly different" (p.405)
is unwarranted.
The
authors said "Bikers traveled at a moderate speed, usually allowing
bicycles to roll down lanes without pedaling where the slope would allow."
Thus it would appear that the mountain biking that they measured is not
representative: it was unusually slow and didn't include much opportunity for
braking, accelerating, or turning, where greater impacts would be expected to
occur.
The
authors also said "Some hikers feel that bikers should be excluded from
existing trails" (p.397). Of course, this is not true. Hikers are only
asking that bikes be excluded, not bikers. On page 407 they admit
the "possibility … that mountain bikers simply contribute further to the
overuse of trails". In other words, allowing bikes on trails allows trail
use to increase over what it would be if bikes weren't allowed. This is
probably true, and deserves to be recognized and researched.
They
found that "One year following treatments, neither vegetation loss nor
species loss was significantly greater on treated lanes than on control
lanes" (p.406). They conclude that the recreation impacts are
"short-term", and experience "rapid recovery". This is
unjustified. Killing plants and destroying seeds modifies the gene pool, and
introduces human-caused loss of genetic diversity, and evolution. Dead plants
and lost genetic diversity do not "recover" (see Vandeman, 2001).
However,
the greatest defect of the study and its interpretation is that is that it
doesn't consider the distance that bikers travel. Even if we accepted their
conclusions that impacts per mile are the same, it would follow that
mountain bikers have several times the impact of hikers, since they are easily
able to, and do, travel several times as far as hikers. Try walking 25
or 50 or 100 miles in a day!
Impacts on Animals:
This
is a very informative paper. The authors "examined the responses of bison
…, mule deer …, and pronghorn antelope … to hikers and mountain bikers … by
comparing alert distance, flight distance, and distance moved" (p.951).
They noted, significantly, that "Outdoor recreation has the potential to
disturb wildlife, resulting in energetic costs, impacts to animals' behavior
and fitness, and avoidance of otherwise suitable habitat. … outdoor
recreation is the second leading cause for the decline of federally threatened
and endangered species on public lands" (p.951). They also noted that
"Mountain biking in particular is one of the fastest-growing outdoor
activities, with 43.3 million persons participating at least once in 2000"
(p.952). However, they didn't draw on this fact when they concluded "We
found no biological justification for managing mountain biking any differently
than hiking" (p.961).
The
authors also surveyed the recreationists, and found that they "failed to
perceive that they were having as great an effect on wildlife as our biological
data indicated. Most recreationists felt that it was acceptable to approach
wildlife at a much closer distance (mean acceptable distance to approach = 59.0
m) than wildlife in our experimental trials would typically allow a human to
approach (mean flight distance of all species = 150.6 m). … Of all visitors
surveyed, 46%, 53%, and 54%, respectively, felt that bison, deer, and pronghorn
were being negatively affected by recreation on
They
noted that the wildlife might habituate to the presence of humans, but that
exactly the opposite happened with the pronghorn: they "in fact used areas
that were significantly farther from trails than they had prior to the start of
recreational use on the island" (p.961). They also noted: "Because
flushing from recreational activity may come at the cost of energy needed for
normal survival, growth, and reproduction …, and because it may cause animals
to avoid otherwise suitable habitat …, it is important that recreationists
understand that their activities can flush wildlife and may make suitable
habitat unavailable" (p.961). I think that the wealth of such information
provided by the authors makes this paper especially valuable.
They
concluded "Our results indicate that there is little difference in
wildlife response to hikers vs. mountain bikers" (p.957). I was present
when Ms. Taylor presented her findings at the Society for Conservation Biology
meeting at the
I
also wonder about the accuracy of their measurements of distance. Distance is
notoriously difficult to measure accurately, especially when animals and
recreationists may be hidden from view ("Due to the inherent errors in
triangulating in the steep canyon country, only ground visual locations were
used in the analysis" p.577). Bias may also have been introduced by the
fact that researchers knew, as they were measuring, which treatment they were
measuring.
Sprung
next cited Papouchis et al (2001), claiming that
"Hikers have [the] greatest impact on bighorn sheep [in
It
is interesting that "when bighorn sheep did respond to human activity,
they noticed vehicles and mountain bikers, on average,
from twice the distance they noticed hikers" (p.577). This would seem to
imply that, were hikers to remain on the trail where the mountain bikers were,
they might have equal or lower impacts than the mountain bikers.
It
is also unfortunate that there was no control group, so that they could
determine the effect of the presence of roads, with and without people on them.
They did note that "avoidance of the road corridor by some animals
represented 15% less use of potential suitable habitat in the high-[visitor-]use area over the low-[visitor-]use area. … human presence in bighorn sheep habitat may cause sheep to
vacate suitable habitat" (p.573). This argues for eliminating all
recreation in the area, especially since the absence of water forces
recreationists to bring motor vehicles carrying water and other supplies:
"mountain bikers frequently use the 161-km White Rim trail, a
4-wheel-drive road. Caravans of mountain bikers accompanied by support vehicles
are common. Day use along the Shafer and White Rim trails exceeded 17,500 vehicles
during the study period, 1993-1994. This use was concentrated from March to
October, with peak use of 134 vehicles/day in May"
(p.575).
The
authors conclude "Contrary to our original expectations and the concerns
of park managers, the increase in numbers of mountain bikers visiting the park
does not appear to be a serious threat to desert bighorn sheep, probably
because mountain bikers are restricted to predictable situations such as the
currently designated road corridors" (p.580). For several reasons, this
conclusion is not justified: (1) as they reported, all recreationists
drive the sheep away from parts of their habitat, causing loss of energy as
well as habitat; (2) permitting bikes causes the total number of visitors to
increase significantly; (3) bikes can't travel alone -- they require motorized
support vehicles, further increasing impacts (e.g. worsening air quality); (4)
there is nothing to prevent mountain bikers from getting off their bikes and
approaching the wildlife; if hikers do that, so will mountain bikers; there is
no reason to exonerate mountain bikers.
They
note, significantly, "However, these results should not be extrapolated to
other public lands where mountain bikers are not confined to designated trails
and may surprise sheep in novel situations" (p.580). Gary Sprung didn't
mention this, thus encouraging inappropriate use of this study's
already-questionable results.
I
would like, however, to commend the authors for stating "we recommend that
park managers manage levels of backcountry activity at low levels"
(p.580). The best policy would be to ban all vehicles, including bicycles (as
well as animals used as vehicles). That would reduce human impacts, without
directly restricting who could go there (perhaps occasional exceptions could be
made for the disabled).
The
authors conclude "Our results show that specific restrictions on mountainbiking above the timberline are not justified from
the point of view of chamois" (p.109). Once again (is there a pattern
here?), this conclusion is not justified. It ignores the fact that mountain
bikers are able to travel several times as far as hikers, and thus negatively
impact several times as much wildlife. It also ignores
the fact that bicycles enable a large increase in numbers of human visitors
(note that this places the blame on the bicycle, not the bicyclists
-- my argument doesn't depend on there being any difference between hikers and
mountain bikers). And, of course, wherever the number of visitors increases,
there is pressure to build more trails, destroying even more habitat. Once
again, it would appear that this study was undertaken with the intent of
excusing mountain biking.
At
the bottom of the same web page is the notice: "IMBA wishes to obtain and
incorporate into future revisions of this document any new or additional
empirical science regarding the impacts of mountain biking. IMBA welcomes
input [my emphasis]. To offer information, please contact the author at
gary@imba.com". On April 25 I emailed Gary (and Pete Webber,
pete@imba.com) the Wisdom et al study, which demonstrates that mountain bikers
have a greater impact on elk than hikers. Not only hasn't this new research
been incorporated into his paper, but I haven't even received a reply. It would
appear that IMBA isn't really interested in achieving a scientific answer to
this question.
In
2003, Jason Lathrop wrote an excellent "critical literature review"
on the ecological impacts of mountain biking, raising some questions found
nowhere else. He quotes the BLM: "An estimated 13.5 million mountain bicyclists visit public lands each year to enjoy the variety
of trails. What was once a low use activity that was easy to manage has become
more complex". He criticizes all of the studies
for not using realistic representations of mountain biking. For example, on
Thurston and Reader, he says "this study's treatment passes at best
loosely approximate the forces exerted by actual mountain biking. On real
trails, riders possess widely varying levels of skill, resulting in variant
speeds, turning, and braking. This study does not address these
variables." Lathrop also makes the excellent point that "Direct
mortality [of animals] is virtually unstudied. I could find no references to it
in the literature. Anecdotal evidence suggests, however, that small mammals are
vulnerable to impact and are not uncommonly killed."
And:
"
Cessford (1995) did an oft-quoted review (which I am including
only because it is so widely cited) that, like all others, uncritically accepts
Finally,
in 2004, Wisdom et al did a very well controlled study comparing the impacts of
ATV riders, mountain bikers, and hikers on elk and mule deer. They say we have
an "urgent need for timely management information to address the rapid
growth in off-road recreation. … Mountain biking [is] … increasing
rapidly". Recreationists were allowed to stop for less than a minute to
look at the animals. All measurements were made electronically, using an
Automated Telemetry System and GPS, allowing control measurements to be made
"blind", with no humans present! "Use of the automated telemetry
system to track animal movements, combined with the use of GPS units to track
human movements, provided real-time, unbiased estimates of the distances
between each ungulate and group of humans [the recreationists were in
pairs]". He pointed out that direct measurements, a la Taylor and Knight, tend to be biased, because some animals
can't be observed. The area was entirely fenced, allowing researchers to
completely control human access.
They
found: "Movement rates of elk were substantially higher during all four
off-road activities as compared to periods of no human activity. … For the
morning pass, movement rates of elk were highest during ATV activity,
second-highest during mountain bike riding, and lowest during hiking and
horseback riding. … Peak movement rates of elk during the morning pass were
highest for ATV riding (21 yards/minute), followed by mountain bike riding (17
yards/minute) and horseback riding and hiking (both about 15 yards/minute). …
By contrast, peak movement rates of elk during the control periods did not
exceed 9 yards/minute during daylight hours of 0800-1500, the comparable period
of each day when off-road treatments were implemented. Interestingly, movement
rates of elk were also higher than control periods at times encompassing
sunrise and sunset for the days in which an off-road activity occurred, even
though humans were not present at these times of the day. These higher movement
rates near sunrise and sunset suggest that elk were displaced from preferred
security and foraging areas as a result of flight behavior during the daytime
off-road activities. In particular, movement rates of elk at or near sunrise
and sunset were higher during the 5-day treatments of mountain bike and ATV
activity".
"Higher
probabilities of flight response occurred during ATV and mountain bike
activity, in contrast to lower probabilities observed during hiking and
horseback riding. Probability of a flight response declined most rapidly during
hiking, with little effect when hikers were beyond 550 yards from an elk. By
contrast, higher probabilities of elk flight continued beyond 820 yards from
horseback riders, and 1,640 yards from mountain bike and ATV riders. In
contrast to elk, mule deer showed less change in movement rates during the four
off-road activities compared to the control periods". (Perhaps they seek
cover, rather than running away.)
"The
energetic costs associated with these treatments deserve further analysis to
assess potential effects on elk survival. For example, if the additional energy
required to flee from an off-road activity reduces the percent body fat below 9
percent as animals enter the winter period, the probability of surviving the
winter is extremely low. Animal energy budgets also may be adversely affected
by the loss of foraging opportunities while responding to off-road activities,
both from increased movements, and from displacement from foraging habitat. …
Our results from 2002 also show clear differences in elk responses to the four
off-road activities. Elk reactions were more pronounced during ATV and mountain
bike riding, and less so during horseback riding and hiking. Both movement
rates and probabilities of flight responses were higher for ATV and mountain
bike riding than for horseback riding and hiking."
It
is also instructive to note that only one pair of ATV users were needed to
cover the 20-mile study area, but two pairs of mountain bikers and three pairs
of hikers were needed, to cover the distance in the time allotted, underscoring
the different relative distances that the three groups are capable of covering.
Summary:
Mountain
bikers have turned to scientific research to try to make mountain biking seem
less harmful, and in particular, to studies comparing it with hiking. Although
they have interpreted this data as indicating that mountain biking impacts are
no greater than those of hiking, a more careful look at these studies leads to
the conclusion that mountain biking impacts are actually several times greater
than those of hikers.
Some
of the important characteristics of mountain biking that have been ignored are:
speed; distance traveled; the increase in number of visitors that bikes allow;
increased trail-building, with its attendant habitat destruction; the displacement
of soil (other than downhill); the killing of roots and soil organisms and
ecosystems; most effects on wildlife; manner of riding (skidding, braking,
acceleration, turning, and representativeness); tire tread; and noise (bikes
are relatively quiet, but a rattling chain may be perceived as
"alien" to natural surroundings).
In
addition, measuring techniques need to be described in more detail,
"blind" measurements should be considered (where the measurers don't
know what treatment they are measuring), controls need to be added, and
"intangibles" (e.g. loss of feelings of safety and loss of the
primitive feel of natural settings) need to be taken more seriously. The direct
killing of small animals deserves attention.
On
the other hand, why do we need research to prove what is obvious? We don't need
any research to know that we shouldn’t step in front of a speeding truck. Or mountain bike.
References:
Cessford, Gordon R. (gcessford@doc.govt.nz), "Off-road impacts
of mountain bikes -- a review and discussion". Science
& Research Series No.92, Department of Conservation, P. O. Box 10-420,
Chiu, Luke (Luke.Chiu@utas.edu.au) and
Lorne Kriwoken (L.K.Kriwoken@utas.edu.au), "Managing
Recreational Mountain Biking in
Foreman, Dave,
"A modest proposal". Wild
Earth, Vol.13, No.1, 2003, pp.34-5, http://www.wildlandsproject.org.
Goeft, Ute and Jackie Alder, "Sustainable mountain biking:
a case study from the southwest of
Hasenauer, Jim (imbajim@aol.com), "A niche for bicycles". Wild Earth,
Vol.13, No.1, 2003, pp.21-22, http://www.wildlandsproject.org.
Lathrop, Jason, "Ecological
impacts of mountain biking: a critical literature review". 2003, http://www.wildlandscpr.org/resourcelibrary/reports/mountainbikingreport.htm.
McCoy, Michael and Mary Alice
Stoner, "Mountain bike trails: Techniques for design, construction and
maintenance". Bikecentennial,
O'Donnell, Brian and Michael
Carroll, "Don't tread here". Wild Earth, Vol.13, No.1, 2003,
pp.31-33, http://www.wildlandsproject.org.
Papouchis, Christopher M. (papouchis@hotmail.com),
Francis J. Singer, and William B. Sloan, "Responses of desert bighorn
sheep to increased human recreation". Journal of Wildlife Management, Vol.65, No.3,
2001, pp.573-82.
Spahr, Robin, "Factors affecting the distribution of bald
eagles and effects of human activity on bald eagles wintering along the
Sprung,
Taylor, Audrey
(audrey_taylor@fws.gov) and Richard L. Knight (knight@cnr.colostate.edu)
"Wildlife responses to recreation and associated visitor
perceptions". Ecological
Applications, Vol.13, No.4, 2003, pp.951-63.
Thurston, Eden and Richard J.
Reader (rreader@uoguelph.ca), "Impacts of experimentally applied mountain
biking and hiking on vegetation and soil of a deciduous forest". Environmental Management, Vol.27, No.3, 2001, pp.397-409.
Vandeman, Michael J. (
Vandeman, Michael J. (
Vandeman, Michael J. (
Wilson, John P. (jpwilson@usc.edu) and Joseph Seney, "Erosional impact of hikers, horses, motorcycles, and off-road
bicycles on mountain trails in
Wisdom, M. J. (mwisdom@fs.fed.us),
Alan A. Ager (aager@fs.fed.us ), H. K. Preisler (hpreisler@fs.fed.us), N. J. Cimon (ncimon@fs.fed.us),
and B. K. Johnson (johnsobd@eou.edu), "Effects of off-road recreation on
mule deer and elk". Transactions of the North
American Wildlife and Natural Resources Conference 69, 2004, pp.531-550.